Garcia's 1-under 71 moved him to 9 under par overall, but his four-shot first-round advantage disappeared as Oosthuizen made an eagle and five birdies in a 67 at Gary Player Country Club.
McIlroy's front nine was promising with three birdies offset by just a single bogey, at the par-four third when his ball plugged just below the lip of a greenside bunker.
Lee, one of 15 players to shoot an opening round 65 in the PGA Tour event, trails American leaders Matt Kuchar, Kramer Hickok and Dominic Bozzelli by a stroke. His round was highlighted by holing a lengthy putt on 17.
It was the Spaniard's lowest opening round score of the season by two shots and matched the biggest first-round lead on the tour in 2018. He ended the day in a tie for 21st.
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The Co Down man returns to Sun City since for the first time since he withdrew with a viral infection at halfway nine years ago, seeking his first win since March and the chance to keep his slim hopes of fourth Race to Dubai crown alive. He began with a birdie and twice reeled off three in a row. "It's never easy, but I've done it before so hopefully I can do it again". "I think I played very smart, I probably wasn't swinging unbelievably but I kept it under control and kept the ball in play all the time which is hard to do here".
"I have to play really well this week to give myself a chance, even half a chance going into next week, and that's what I'm going to try and do", said McIlroy, who is eighth in the Race to Dubai, nearly two million points behind Molinari. I rolled the ball nicely, I made some good putts here and there. "I kept the ball in play all the time, which is hard to do here".
Garcia won a third consecutive Andalucia Valderrama Masters title last time out after another strong showing in Europe's Ryder Cup triumph.
Scrivener also started well at last week's Turkish Airlines Open before a perhaps disappointing weekend saw him finish 23rd but with the carrot of a start at next week's season ending DP World Tour Championship in Dubai still available to him with a good finish this week he has a lot still to play for.