Lawmakers will vote on Wednesday on whether to dismiss a plan proposed by the upper chamber of parliament which would require ministers to report what efforts they had made to secure a customs union with the EU by the end of October.
Frank Field, a veteran lawmaker from the northwest of England, was his first victim after suggesting the point of the amendment Pennycook was backing was to stop Brexit, then Gareth Snell, who represents Stoke-on-Trent in the Midlands, got the same treatment for making a similar point. He said he would vote against the prime minister.
Dominic Grieve withdrew his own amendment, which would have given MPs powers to dictate what the Government should do if no acceptable agreement is reached by February 2019.
But it was a vote in parliament on Tuesday that left her seemingly at the mercy of two groups in the Conservative Party - those who want to maintain the closest possible ties with the European Union, and others pressing for a clean break. More than half of Leave voters (57%) said they too would be concerned to have fewer rights and protections after Brexit. Pro-Brexit Conservative lawmaker Edward Leigh slammed pro-EU colleagues, saying Parliament must respect the result of the June 2016 voter referendum.
Mr Grieve confirmed he voted with the UK Government after receiving assurances, telling Sky News: "I am quite satisfied we are going to get a meaningful vote".
A spokesman for Ms MacLean said she had never spoken to Best for Britain or attended an event held by them, describing suggestions she could resign over Brexit as "utter rubbish".
In an effort to bring some clarity to the debate - and to dispel allegations made by some in the United Kingdom, including the government's own chief Brexit negotiator, that Brussels is being hard in negotiations to punish Britain for leaving - European Union figures have become very fond of going back to Brexit basics.
The defeated amendments, from the House of Lords, include one handing Parliament the final say on any deal the government strikes with Brussels.
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To avoid defeat, the government promised to make its own changes to the bill to strengthen Parliament's powers.
But pro-EU Tories are warning that they remain ready to rebel if their demands are not satisfied by the compromise amendment, expected to be tabled on Thursday ahead of the bill's return to the Lords on Monday.
The Brexit talks are progressing painfully slowly, but both sides still hope to reach a deal in October, ahead of Britain's withdrawal from the European Union in March 2019. The difference in this particular case is the enormity of the issue in the context of Brexit and in a climate of suspicion and hostility from many on the Conservative backbenches towards their leader.
Yet this week's marathon Brexit debate have Labour members of Parliament concerned about their Brexit-voting electorates - and pro-European lawmakers frustrated by Corbyn's euroskeptic history - airing frustrations in full view.
But the resignation by Phillip Lee, who has always been critical of the government's Brexit strategy, underlined the deep rifts in the party over Brexit that makes such votes anything but easy.
The concession came after intensive horse-trading on the floor of the House of Commons, with chief whip Julian Smith shuttling between Tory backbenchers during debate on Lords amendments to the EU Withdrawal Bill.
It has also intensified pressure on a prime minister who lost her party's parliamentary majority at an ill-judged election past year and tested her already weakened authority. She had nearly lost a general election she never needed to call, squandering her Conservative Party's overall majority and forcing her to rely on the support of the smaller Democratic Unionist Party.
The Government has not and will not agree to MPs binding its hands in the Brexit negotiations, officials have insisted, after the Prime Minister saw off a threatened rebellion with the promise of concessions.